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Part of this may be due to the difficulty defining "normal" testosterone levels and "normal" behavior. of testosterone vary dramatically over time and even during the course of a day. In addition, what may seem like a symptom of testosterone excess may actually be unrelated to this hormone. Testosterone and the classical nuclear androgen receptor first appeared in gnathostomes . Agnathans such as lampreys do not produce testosterone but instead use androstenedione as a male sex hormone. Both testosterone and 5α-DHT are metabolized mainly in the li